Link to Google Scholar Page

Roussel, A, AA Pavlov, J Dworkin, and SS Johnson, Rapid destruction of lipid biomarkers under simulated cosmic radiation, in press, Astrobiology.

Roussel, AR, A McAdam, A Pavlov, C Knudson, C Achilles, D Foustoukos, J Dworkin, and SS Johnson, Variable and Large Losses of Diagnostic Biomarkers After Simulated Cosmic Radiation Exposure in Clay- and Carbonate-Rich Mars Analog Samples, 2024, Astrobiology, 24(7):669-683,  doi: 10.1089/ast.2023.0123.

Fishman, CB, JG Bevilacqua, AS Hahn, C Morgan-Lang, N Wagner, O Gadson, AC McAdam, J Bleacher, C Achilles, C Knudson, MM Millan, DM Bower, M Musilova, and SS Johnson, Extreme Niche Partitioning and Microbial Dark Matter in a Mauna Loa Lava Tube, 2023, Journal of Geophysical Research, e2022JE007283.

Azua-Bustos, A, A Farien, C Gonzales-Silva, O Preito-Ballesteros, D Carrizo, L Zanchez-Garcia, V Parro, MA Fernandez-Martinez, C Escudero, V Munoz-Iglesias, M Fernandez-Sampedro, A Molina, M Garcis-Villadangos, M Moreno-Paz, J Wierzchos, A Ascaso, T Fornaro, J Brucato, G Poggiali, JA Manrique, M Veneranda, G Lpez-Reyes, A Zanz-Arranz, F Rull, A Ollila, R Wiens, A Reyes-Newll, S. Clegg, M Millan, SS Johnson, O McIntosh, C Szopa, C Freissinet, Y Sekine, K Fukishi, K Morida, K Inoue, H Sakuma, and E Rampe, Dark microbiome and extremely low organics in an Atacama fossil river delta unveil Mars life detection limits, 2023, Nature Communications, 14(808), doi.org/10.1038/s41467-023-36172-1.

Bower, DM, AC McAdam, CSC Yang, M Millan, R Arevalo, C Achilles, C Knudson, T Hewagama, CA Nixon, CB Fishman, SS Johnson, J Bleacher, P Whelley, Spectroscopic comparisons of two different terrestrial basaltic environments: Exploring the correlation between nitrogen compounds and biomolecular signatures, 2023, Icarus, doi.org/10.1016/j.icarus.2023.115626. 

Fishman, CB, JG Bevilacqua, O Gadson, AS Hahn, AC McAdam, J. Bleacher, SS Johnson, Basaltic lava tubes host a putative novel genus in the family Solirubrobacteraceae, 2022, Microbiology Resource Announcements, 11(10):e0049922, doi.org/10.1128/mra.00499-22.

Millan, M, AJ Williams, AC Mcadam, JL Eigenbrode, A Steele, C Freissinet, DP Glavin, C Szopa, A Buch, RE Summons, JMT Lewis, GM Wong, CH House, B Sutter, O Mcintosh, AB Bryk, HB Franz, C Pozarycki, JC Stern, R Navarro‐Gonzalez, DP Archer, V Fox, K Bennett, S Teinturier, C Malespin, SS Johnson and P Mahaffy, Sedimentary Organics in Glen Torridon, Gale Crater, Mars: Results from the SAM Instrument Suite and Supporting Laboratory Analyses, 2022, Journal of Geophysical Research: Planets (Special Issue), e2021JE007107, doi.org/10.1029/2021JE007107.

Gadson O, JG Bevilacqua, Fishman, CB, AS Hahn, AC McAdam, J. Bleacher, SS Johnson, Metagenome-assembled genome of a putative chemoheterotroph from volcanic terrain in Hawaii, 2022, Microbiology Resource Announcements, doi: 10.1128/mra.00556-22.

Michalski, JR, TA Goudge, S Crowe, J Cuadros, J Mustard, and SS Johnson, 2022, Geological Diversity and Microbiological Potential of Lakes on Mars, Nature Astronomy, doi.org/10.1038/s41550-022-01743-7.

Weng, M.M., E Zaikova, M Millan, AJ Williams, AC McAdam, CA Knudson, SR Fuqua, NY Wagner, K Craft, S Kobs Nawotniak, A Shields, J Bevilacqua, Y Bai, SS Hughes, WB Garry, JL Heldmann, DSS Lim, D Buckner, P Gant, SS Johnson, Life Underground: Investigating Microbial Communities and their Biomarkers in Mars-analog Lava Tubes at Craters of the Moon National Monument and Preserve, 2022, Journal of Geophysical Research, 127 (11), e2022JE007268, doi.org/10.1029/2022JE007268.

Roussel, AR, AC McAdam, HV Graham, AA Pavlov, CN Achilles, CA Knudson, A Steele, DI Foustoukos, and SS Johnson, Diagnostic Biosignature Transformation Under Simulated Martian Radiation in Organic-rich Sedimentary Rocks, 2022, Frontiers in Astronomy and Space Sciences, 9:919828, doi.org/10.3389/fspas.2022.919828.

McAdam, AC, B Sutter, PD Archer, HB Franz, GM Wong, JM Lewis, JV Clark, M Millan, AJ Williams, JL Eigenbrode, CA Knudson, C Freissinet, DP Glavin, JC Stern, R Navarro‐González, CN Achilles, DW Ming, RV Morris, TF Bristow, EB Rampe, MT Thorpe, CH House, S Andrejkovičová, AB Bryk, VK Fox, SS Johnson, PR Mahaffy, and CA Malespin, 2022. Evolved gas analyses of sedimentary rocks from the Glen Torridon Clay‐Bearing Unit, Gale crater, Mars: Results from the Mars Science Laboratory Sample Analysis at Mars Instrument Suite. Journal of Geophysical Research: Planets, e2022JE007179.

Wagner, N, D Andersen, A.Hahn, and SS Johnson, 2022. Metagenomics of an Anoxic Microbial Ecosystem in Lake Untersee, a Potential Analog for Enceladus, Scientific Reports.12, 7376. doi.org/10.1038/s41598-022-10876-8.

Millan, M, S Teinturier, CA Malespin, JY Bonnet, A Buch, JP Dworkin, JL Eigenbrode, C Freissinet, DP Glavin, R Navarro-Gonzalez, A Srivastava, JC Stern, B Sutter, C Szopa, AJ Williams, RH Williams, GW Wong, SS Johnson and P Mahaffy, 2022. Organic molecules revealed in Mars’s Bagnold Dunes by Curiosity’s derivatization experiment. Nature Astronomy, pp.1-12. doi.org/10.1038/s41550-021-01507-9.

Wagner, NY, D Andersen, A Hahn, R McLaughlin, SS Johnson, 2022. Draft Genome Sequence from a Putative New Genus and Species in the Family M1A02 within Planctomycetes, Isolated from Benthic Pinnacle Mats of Lake Untersee, Antarctica, Microbiology Resource Announcements, doi.org/10.1128/mra.01192-21.

Chou, L, P Mahaffy, M Trainer, J Eigenbrode, R Arevalo, W Brinckerhoff, S Getty, N Grefenstette, V Da-Poian, G Fricke, C Kempes, J Marlow, B Sherwood Lollar, H Graham, and SS Johnson, 2021. Planetary Mass Spectrometry for Agnostic Life Detection in the Solar System. Frontiers in Astronomy and Space Sciences. 755100. doi.org/10.3389/fspas.2021.755100.

Smith, HH, AS Hyde, DN Simkus, E Libby, SE Maurer, HV Graham, CP Kempes, B Sherwood Lollar, L Chou, AD Ellington, GM Fricke, PR Girguis, NM Grefenstette, CI Pozarycki, CH House, and SS Johnson, 2021. The Grayness of the Origin of Life. Life. 11(6):498. doi.org/ 10.3390/life11060498.

Williams, A, KL Craft, M Millan, SS Johnson, CA Knudson, MJ Rivera AC McAdam, D Tobler, and JR Skok, 2021, Fatty Acid Preservation in Modern and Relict Hot Spring Deposits in Iceland, with Implications for Organics Detection on Mars. Astrobiology,  21(1), pp.60-82.21(1), doi.org/10.1089/ast.2019.2115.

He, Y, A Buch, C Szopa, M Millan, C Freissinet, R Navarro-Gonzalez, M Guzman, SS Johnson, D Glavin, A Williams, and J Eigenbrode, 2021. Influence of Calcium Perchlorate on the Search for Martian Organic Compounds with MTBSTFA/DMF Derivatization. Astrobiology, 21(9), pp.1137-1156. doi.org/10.1089/ast.2020.2252.

Chou, L, N Grefenstette, SS Johnson, H Graham, P Mahaffy, C Kempes, … BS Lollar, 2021. Towards a more universal life detection strategy. Bulletin of the AAS, 53(4). doi.org/10.3847/25c2cfeb.53a24171.

Banfield, D, J Stern, A Davila, SS Johnson, D Brain, R Wordsworth, … K Watts, 2021. Summary of the Mars Science Goals, Objectives, Investigations, and Priorities. Bulletin of the AAS, 53(4). doi.org/10.3847/25c2cfeb.4548c524.

Schmidt, B, SS Johnson, T Hoehler, H Graham, J Bowman, S Som, … J Buffo, 2021. Enabling Progress Towards Life Detection on NASA Missions. Bulletin of the AAS, 53(4). doi.org/10.3847/25c2cfeb.77a5ad8e.

Johnson, SS, HV Graham, DJ Des Marais, and RM Hazen, 2020. Detecting Life on Earth and the Limits of Analogy. In Planetary Astrobiology (V. Meadows et al., eds.), pp. 121-150. University of Arizona Press. | PDF

Millan, M, C Szopa, A Buch, RE Summons, R Navarro‐Gonzalez, P Mahaffy, and SS Johnson, 2020. Influence of Calcium Perchlorate on Organics under SAM-like Pyrolysis Conditions: Constraints on the Nature of Martian Organics. Journal of Geophysical Research: Planets, 125(7), e2019JE006359. doi.org/10.1029/2019JE006359 | PDF

Johnson, SS, M Millan, H Graham, KC Benison, AJ Williams, A McAdam, CA Knudson, S Andrejkovičová, and C Achilles, 2020. Lipid Biomarkers in Ephemeral Acid Salt Lake Mudflat/Sandflat Sediments: Implications for Mars, Astrobiology, 20(2), 167-178. doi.org/10.1089/ast.2017.1812. | PDF

Fraeman, AA, LA Edgar, EB Rampe, LM Thompson, J Frydenvang, CM Fedo, JG Catalano, WE Dietrich, TSJ Gabriel, AR Vasavada, JP Grotzinger, J L’Haridon, N Mangold, VZ Sun, CH House, AB Bryk, C Hardgrove, S Czarnecki, KM Stack, RV Morris, RE Arvidson, SG Banham, KA Bennett, JC Bridges, CS Edwards, WW Fischer, VK Fox, S Gupta, BHN Horgan, SR Jacob, JR Johnson, SS Johnson, DM Rubin, MR Salvatore, SP Schwenzer, KL Siebach, NT Stein, S Turner, DF Wellington, RC Wiens, AJ Williams, G David, and GM Wong, 2020. Evidence for a Diagenetic Origin of Vera Rubin Ridge, Gale Crater, Mars: Summary and Synthesis of Curiosity’s Exploration Campaign. Journal of Geophysical Research: Planets, e2020JE006527.doi.org/10.1029/2020JE006527.

Greenfield, SR, SW Tighe, Y Bai, DS Goerlitz, M Von Turkovich, DJ Taatjes, JA Dragon, and SS Johnson, 2020. Life and its Traces in Antarctica’s McMurdo Dry Valley Paleolakes: A Survey of Preservation, Micron, 131. doi.org/10.1016/j.micron.2019.102818. | PDF

Buch, A, I Belmahdi, C Szopa, C Freissinet, DP Glavin, M Millan, R Summons, D Coscia, S Teinturier, J‐Y Bonnet, Y He, M Cabane, R Navarro‐Gonzalez, CA Malespin, J Stern, J Eigenbrode, PR Mahaffy, and SS Johnson, 2019. Role of the Tenax Adsorbent in the Interpretation of the EGA and GC‐MS Analyses Performed With the Sample Analysis at Mars in Gale Crater, Journal of Geophysical Research: Planets, 124, 2819-2851. doi.org/10.1029/2019JE005973. | PDF

Wagner, NY, AS Hahn, D Andersen, MB Wilhelm, C Morgan-Lang, M Vanderwilt, and SS Johnson, 2019. Draft Genome Sequence from a Putative New Genus and Species in the Family Methanoregulaceae Isolated from the Anoxic Basin of Lake Untersee in East Antarctica, Microbiology Resource Announcements, 8(18): e00271-19. doi.org/10.1128/MRA.00271-19. | PDF

Millan, M, C Szopa, A Buch, M Cabane, S Teinturier, P Mahaffy, and SS Johnson, 2019. Performance of the SAM Gas Chromatographic Columns under Simulated Flight Operating Conditions for the Analysis of Chlorohydrocarbons on Mars, Journal of Chromatography A, 1598, 183-195. doi.org/10.1016/j.chroma.2019.03.064. | PDF

Williams, AJ, J Eigenbrode, M Floyd, MB Wilhelm, S O’Reilly, SS Johnson, KL Craft, CA Knudson, S Andrejkovičová, JMT Lewis, A Buch, DP Glavin, C Freissinet, RH Williams, C Szopa, M Millan, RE Summons, A McAdam, K Benison, R Navarro-González, C Malespin, and PR Mahaffy, 2019. Recovery of Fatty Acids from Mineralogic Mars Analogs by TMAH Thermochemolysis for the Sample Analysis at Mars Wet Chemistry Experiment on the Curiosity Rover, Astrobiology, 19(4): 522-546. doi.org/10.1089/ast.2018.1819. | PDF

Sutton, MA, AS Burton, E Zaikova, RE Sutton, WB Brinckerhoff, JG Bevilacqua, MM Weng, MJ Mumma, and SS Johnson, 2019. Radiation Tolerance of Nanopore Sequencing Technology for Life Detection on Mars and Europa, Scientific Reports, 9: 5370. doi.org/10.1038/s41598-019-41488-4. | PDF

He, Y, A Buch, M Morisson, C Szopa, C Freissinet, A Williams, M Millan, M Guzman, R Navarro-Gonzalez, J-Y Bonnet, D Coscia, JL Eigenbrode, CA Malespin, P Mahaffy, DP Glavin, JP Dworkin, P Lu, and SS Johnson, 2019. Application of TMAH Thermochemolysis to the Detection of Nucleobases: Application to the MOMA and SAM Space Experiment, Talanta, 204, 802-811. doi.org/10.1016/j.talanta.2019.06.076. | PDF

Zaikova, E, DS Goerlitz, SW Tighe, NY Wagner, Y Bai, BL Hall, JG Bevilacqua, MM Weng, MD Samuels-Fair, and SS Johnson, 2019. Antarctic Relic Microbial Mat Community Revealed by Metagenomics and Metatranscriptomics, Frontiers in Ecology and Evolution, doi.org/10.3389/fevo.2019.00001. | PDF

Michalski, J, T Glotch, AD Rogers, P Niles, J Cuadros, J Ashley, and SS Johnson, 2019. The geology and astrobiology of McLaughlin Crater, Mars: an ancient lacustrine basin containing turbidites, mudstones and serpentinites, Journal of Geophysical Research, doi.org/10.1029/2018JE005796. | PDF

Sun, VZ, KM Stack, LC Kah, L Thompson, W Fischer, AJ Williams, SS Johnson, RC Wiens, RE Kronyak, M Nachon, CH House, and S VanBommel, 2018. Late-Stage Diagenetic Concretions in the Murray Formation, Gale Crater, Mars, Icarus, doi.org/10.1016/j.icarus.2018.12.030. | PDF

Johnson, SS, E Anslyn, H Graham, P Mahaffy, and A Ellington, 2018. Fingerprinting Non-Terran Life, Astrobiology, doi.org/10.1089/ast.2017.1712. | PDF

Zaikova, E, K Benison, M Mormile, and SS Johnson, 2018. Microbial communities and their predicted metabolic functions in a desiccating acid salt lake, Extremophiles, 22(3), 367-379. doi.org/10.1007/s00792-018-1000-4. | PDF

Johnson, SS, H Graham, E Anslyn, P Conrad, L Cronin, A Ellington, J Elsila, P Girguis, C House, C Kempes, E Libby, P Mahaffy, J Nadeau, MR Rwebangirai, B Sherwood Lollar, and A Steele, 2018. Agnostic Biosignatures: Toward a More Inclusive Life Detection Strategy, White Paper for the National Academies Committee on the Astrobiology Science Strategy for the Search for Life in the Universe | PDF

Ehlmann, B, E Kite, D Rogers, T Glotch, C Fassett, A Hayes, C Edwards, J Mustard, A Fraeman, K Stack, B Horgan, PB Niles, L Kerber, M Rice, and SS Johnson, 2018. Mars as a Linchpin for the Understanding the Habitability of Terrestrial Planets: Discoveries of the Last Decade from Mars and Why A New Paradigm of Multiple, Landed Robotic Explorers is Required for Future Progress in Terrestrial Planet Astrobiology, White Paper for the National Academies Committee on the Astrobiology Science Strategy for the Search for Life in the Universe | PDF

Michalski, JR, TC Onstott, SJ Mojzsis, J Mustard, Q Chan, PB Niles, and SS Johnson, 2017. The Martian subsurface as a potential window into the origin of life, Nature Geoscience, 11, 21-26. doi.org/10.1038/s41561-017-0015-2. | PDF

Franz, H, A McAdam, DW Ming, C Freissinet, P Mahaffy, D Eldridge, W Fischer, J Grotzinger, C House, J Hurowitz, S McLennan, S Schwenzer, D Vaniman, P Archer, S Atreya, P Conrad, J Dottin, J Eigenbrode, K Farley, Daniel Glavin, SS Johnson, C Knudson, R Morris, R Navarro-Gonzalez, A Pavlov, R Plummer, E Rampe, J Stern, A Steele, R Summons, and B Sutter, 2017.  Large Sulphur Isotope Fractionations in Martian Sediments at Gale Crater. Nature Geoscience, 10(9), 658-662. doi:10.1038/ngeo3002. | PDF

Johnson, SS, E Zaikova, D Goerlitz, Y Bai, and S Tighe, 2017. Real Time DNA Sequencing in the Antarctic Dry Valleys using the Oxford Nanopore Sequencer. Journal of Biomolecular Techniques, 28(1), 2-7, doi: 10.7171/jbt.17-2801-009. | PDF

Tighe, S, E Afshinnekoo, TM Rock, K McGrath, N Alexander, A McIntyre, S Ahsanuddin, D Bezdan, SJ Green, S Joye, SS Johnson, DA Baldwin, N Bivens, N Ajami, JR Carmical, IC Herriott, R Colwell, M Donia, J Foox, N Greenfield, T Hunter, J Hoffman, J Hyman, E Jorgensen, D Krawczyk, J Lee, S Levy, N Garcia-Reyero, M Settles, K Thomas, F Gómez, L Schriml, N Kyrpides, E Zaikova, J Penterman, and CE Mason, 2017. Genomic Methods and Microbiological Technologies for Profiling Novel and Extreme Environments for the Extreme Microbiome Project (XMP). Journal of Biomolecular Techniques, 28(1): 31–39. doi: 10.7171/jbt.17-2801-004. | PDF

Johnson, SS, MG Chevrette, BL Ehlmann, and K Benison, 2015. Insights from the metagenome of an acid salt lake: the role of biology in an extreme depositional environment. PLOS One, 10(4). doi: 10.1371/journal.pone.0122869. | PDF

Johnson, SS, AA Pavlov, and MA Mischna, 2009. Fate of SO2 in the ancient Martian atmosphere: implications for transient greenhouse warming. Journal of Geophysical Research, 114, E11011. doi:10.1029/2008JE003313. | PDF

Johnson, SS, MA Mischna, TL Grove, and MT Zuber, 2008. Sulfur-induced Greenhouse Warming on Early Mars. Journal of Geophysical Research, 113, E08005. doi: 10.1029/2007JE002962. | PDF

Isenbarger, TA, CE Carr, SS Johnson, M Finney, GM Church, W Gilbert, MT Zuber, and G Ruvkun, 2008. The most conserved genome segments for life detection on Earth and other planets. Origins of Life and Evolution of Biospheres, 38(6), 517–533. doi:10.1007/s11084-008-9148-z. | PDF

Johnson, SS, MB Hebsgaard, TR Christensen, M Mastepanov, R Nielsen, K Munch, T Brand, MTP Gilbert, MT Zuber, M Bunce, R Rønn, D Gilichinsky, D Froese, and E Willerslev, 2007. Ancient Bacteria Show Evidence of DNA Repair. Proceedings of the National Academy of Sciences, 104(36), 14401-14405. doi: 10.1073/pnas.0706787104. | PDF

Herkenhoff, KE, SW Squyres, R Anderson, BA Archinal, RE Arvidson, JM Barrett, KJ Becker, JF Bell, C Budney, NA Cabrol, MG Chapman, D Cook, BL Ehlmann, J Farmer, B Franklin, LR Gaddis, DM Galuszka, PA Garcia, TM Hare, E Howington-Kraus, JR Johnson, SS Johnson, K Kinch, RL Kirk, EM Lee, C Leff, M Lemmon, MB Madsen, JN Maki, KF Mullins, BL Redding, L Richter, MR Rosiek, MH Sims, LA Soderblom, N Spanovich, R Springer, RM Sucharski, T. Sucharski, R Sullivan, JM Torson, and A Yen, 2006, Overview of the Microscopic Imager Investigation during Spirit’s first 450 sols in Gusev crater. Journal of Geophysical Research. 111, E02S04, doi:10.1029/2005JE002574. | PDF

McLennan, SM, JF Bell III, WM Calvin, PR Christensen, BC Clark, PA de Souza, JD Farmer, WH Farrand, DA Fike, R Gellert, A Ghosh, TD Glotch, JP Grotzinger, B Hahn, KE Herkenhoff, J. A Hurowitz, JR Johnson, SS Johnson, BL Joliff, G Klingelhöfer, AH Knoll, Z Learner, MC Malin, HY McSween Jr., J Pocock, SW Ruff, LA Soderblom, SW Squyres, NJ Tosca, WA Watters, MB Wyatt, and A Yen, 2005. Provenance and diagenesis of the evaporate-bearing Burns formation, Meridiani Planum, Mars. Earth and Planetary Science Letters, 240, 95 – 121. doi: 10.1016/j.epsl.2005.09.041 | PDF

Ehlmann, BL, RE Arvidson, BL Jolliff, SS Johnson, B Ebel, N Lovenduski, JD Morris, JA Byers, NO Snider, and RE Criss, 2005. Hydrologic and Isotopic Modeling of Alpine Lake Waiau. Pacific Science, 59(1), 1-15. doi: 10.1353/psc.2005.0005. | PDF